The world champions, who had not lost to England for four years, rarely looked in danger of losing that proud record as they ran in tries from flanker Willem Alberts and wing Lwazi Mvovo to win a bruising contest.
Victory was no less than South Africa deserved after a ferociously committed effort that left England battered -- both flanker Tom Croft and fly-half Flood departed injured long before the final whistle.
England's captain Lewis Moody described the defeat as "bitterly disappointing."
"Fair play to South Africa -- they played exceptionally well today. We're bitterly disappointed," Moody said.
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